# Lake Ohrid DEEP (Albania/N. Macedonia) Biomarker, Stable Isotope, Diatom and Pollen Data from 1365-1165 ka #----------------------------------------------------------------------- # World Data Service for Paleoclimatology, Boulder # and # NOAA Paleoclimatology Program #----------------------------------------------------------------------- # Template Version 3.0 # Encoding: UTF-8 # NOTE: Please cite original publication, online resource and date accessed when using this data. # If there is no publication information, please cite Investigator, title, online resource and date accessed. # # Description/Documentation lines begin with # # Data lines have no # # # Online_Resource: https://www.ncdc.noaa.gov/paleo/study/28070 # Description: NOAA Landing Page # Online_Resource: https://www1.ncdc.noaa.gov/pub/data/paleo/paleolimnology/europe/southern_europe/panagiotopoulos2019/panagiotopoulos2019-pollen.txt # Description: NOAA location of the template # # Data_Type: Paleolimnology # # Dataset_DOI: # # Parameter_Keywords: geochemistry, physical properties, oxygen isotopes, carbon isotopes, population abundance #--------------------------------------- # Contribution_Date # Date: 2019-11-22 #--------------------------------------- # File_Last_Modified_Date # Date: 2019-11-22 #--------------------------------------- # Title # Study_Name: Lake Ohrid DEEP (Albania/N. Macedonia) Biomarker, Stable Isotope, Diatom and Pollen Data from 1365-1165 ka #--------------------------------------- # Investigators # Investigators: Panagiotopoulos, Konstantinos #--------------------------------------- # Description_Notes_and_Keywords # Description: Further details available in the publication: https://www.sciencedirect.com/science/article/pii/S0277379119307127?via%3Dihub # Provided Keywords: Mediterranean region, mid-altitude refugium, relicts, vegetation succession, pollen-based, biome reconstruction, palynological richness, microscopic charcoal, lipid biomarkers, diatoms, stable isotopes #--------------------------------------- # Publication # Authors: Panagiotopoulos, K., Holtvoeth, J., Kouli, K., Marinova, E., Francke, A., Cvetkoska, A., Jovanovska, E., Lacey, J., Lyons, E. T., Buckel, C., Bertini, A., Donders, T., Just, J., Leicher, N., Leng, M. J., Melles, M., Pancost, R., Sadori, L., Tauber, P., Vogel, H., Wagner, B., and Wilke, T. # Published_Date_or_Year: 2019 # Published_Title: Insights into the evolution of the young Lake Ohrid ecosystem and vegetation succession from a southern European refugium during the Early Pleistocene # Journal_Name: Quaternary Science Reviews # Volume: # Edition: # Issue: # Pages: # Report_Number: # DOI: 10.1016/j.quascirev.2019.106044 # Online_Resource: # Full_Citation: # Abstract: Mediterranean mid-altitude sites are critical for the survival of plant species allowing for elevational vegetation shifts in response to high-amplitude climate variability. Pollen records from the southern Balkans have underlined the importance of the region in preserving plant diversity over at least the last half a million years. So far, there are no records of vegetation and climate dynamics from Balkan refugia with an Early Pleistocene age. Here we present a unique palynological archive from such a refugium, the Lake Ohrid basin, recording continuously floristic diversity and vegetation succession under obliquity-paced climate oscillations. Palynological data are complemented by biomarker, diatom, carbonate isotope and sedimentological data to identify the mechanisms controlling shifts in the aquatic and terrestrial ecosystems within the lake and its catchment. The study interval encompasses four complete glacial-interglacial cycles (1365-1165 ka; MIS 43-35). Within the first 100 kyr of lake ontogeny, lake size and depth increase before the lake system enters a new equilibrium state as observed in a distinct shift in biotic communities and sediment composition. Several relict tree genera such as Cedrus, Tsuga, Carya, and Pterocarya played an important role in ecological succession cycles, while total relict abundance accounts for up to half of the total arboreal vegetation. The most prominent biome during interglacials is cool mixed evergreen needle leaf and deciduous broadleaf forests, while cool evergreen needleleaf forests dominate within glacials. A rather forestedlandscape with a remarkable plant diversity provide unique insights into Early Pleistocene ecosystem resilience and vegetation dynamics. #--------------------------------------- # Funding_Agency # Funding_Agency_Name: German Research Foundation (DFG) # Grant: PA 2664/2-1 #--------------------------------------- # Site_Information # Site_Name: Lake Ohrid ICDP5045-1 # Location: Southern Europe # Country: Albania/North Macedonia # Northernmost_Latitude: 41.04917 # Southernmost_Latitude: 41.04917 # Easternmost_Longitude: 20.715 # Westernmost_Longitude: 20.715 # Elevation: 693 #--------------------------------------- # Data_Collection # Collection_Name: Lake Ohrid DEEP(5045-1) Palynology Panagiotopoulos2019 # First_Year: 1365000 # Last_Year: 1165000 # Time_Unit: cal yr BP # Core_Length: 447.28-400.00 # Notes: #--------------------------------------- # Chronology_Information # Chronology: #--------------------------------------- # Variables # Data variables follow that are preceded by "##" in columns one and two. # Variables list, one per line, shortname-tab-longname components (9 components: what, material, error, units, seasonality, archive, detail, method, C or N for Character or Numeric data) ## age_ka age,,,calendar kiloyear before present,,paleolimnology,,,N, ## depth_m depth,,,meter,,paleolimnology,,,N, ## Pollen_Sum Pollen Sum (counts),,,count,,paleolimnology,,,N,Pollen Sum (counts) ## Pollen_Sum-pine Pollen Sum - Pines (counts),,,count,,paleolimnology,,,N,Pollen Sum - Pines (counts) ## Relict% Relict %,,,percent,,paleolimnology,,,N,Relict % ## Mesophilous% Mesophilous %,,,percent,,paleolimnology,,,N,Mesophilous % ## Mesophilous%-pine Mesophilous % (- Pines),,,percent,,paleolimnology,,,N,Mesophilous % (- Pines) ## Mesophilous_Conc Mesophilous Conc (grains/g),,,grain per gram,,paleolimnology,,,N,Mesophilous Conc (grains/g) ## Montane% Montane %,,,percent,,paleolimnology,,,N,Montane % ## Montane%-pine Montane % (- Pines),,,percent,,paleolimnology,,,N,Montane % (- Pines) ## Montane_Conc Montane Conc (grains/g),,,grain per gram,,paleolimnology,,,N,Montane Conc (grains/g) ## Pines% Pines %,,,percent,,paleolimnology,,,N,Pines % ## Pines_Conc Pines Conc (grains/g),,,grain per gram,,paleolimnology,,,N,Pines Conc (grains/g) ## AP% AP %,,,percent,,paleolimnology,,,N,AP % ## AP%-pine AP % (- Pines),,,percent,,paleolimnology,,,N,AP % (- Pines) ## AP_Conc AP Conc (grains/g),,,grain per gram,,paleolimnology,,,N,AP Conc (grains/g) ## NAP% NAP %,,,percent,,paleolimnology,,,N,NAP % ## NAP%-pine NAP % (- Pines),,,percent,,paleolimnology,,,N,NAP % (- Pines) ## NAP_Conc NAP Conc (grains/g),,,grain per gram,,paleolimnology,,,N,NAP Conc (grains/g) ## Poaceae% Poaceae %,,,percent,,paleolimnology,,,N,Poaceae % ## Poaceae%-pine Poaceae % (- Pines),,,percent,,paleolimnology,,,N,Poaceae % (- Pines) ## Poaceae_Conc Poaceae Conc (grains/g),,,grain per gram,,paleolimnology,,,N,Poaceae Conc (grains/g) ## Aquatics% Aquatics %,,,percent,,paleolimnology,,,N,Aquatics % ## Aquatics%-pine Aquatics % (- Pines),,,percent,,paleolimnology,,,N,Aquatics % (- Pines) ## Aquatics_Conc Aquatics Conc (grains/g),,,grain per gram,,paleolimnology,,,N,Aquatics Conc (grains/g) ## Charcoal_Conc Charcoal Conc (particles/g),,,particle per gram,,paleolimnology,,,N,Charcoal Conc (particles/g) ## Pediastrum_Conc Pediastrum Conc (coenobia/g),,,coenobia per gram,,paleolimnology,,,N,Pediastrum Conc (coenobia/g) ## Botryococcus_Conc Botryococcus Conc (coenobia/g),,,coenobia per gram,,paleolimnology,,,N,Botryococcus Conc (coenobia/g) ## E-T400-min E(T400) min,,,,,paleolimnology,,,N,E(T400) min ## E-T400-mean E(T400) mean,,,,,paleolimnology,,,N,E(T400) mean ## E-T400-max E(T400) max,,,,,paleolimnology,,,N,E(T400) max ## pred_biome0.5%_thresh notes,,,,,paleolimnology,,,C,Predicted Biome_0.5% threshold #------------------------ # Data: # Data lines follow (have no #) # Data line format - tab-delimited text, variable short name as header # Missing_Values: -999 age_ka depth_m Pollen_Sum Pollen_Sum-pine Relict% Mesophilous% Mesophilous%-pine Mesophilous_Conc Montane% Montane%-pine Montane_Conc Pines% Pines_Conc AP% AP%-pine AP_Conc NAP% NAP%-pine NAP_Conc Poaceae% Poaceae%-pine Poaceae_Conc Aquatics% Aquatics%-pine Aquatics_Conc Charcoal_Conc Pediastrum_Conc Botryococcus_Conc E-T400-min E-T400-mean E-T400-max pred_biome0.5%_thresh 1164.71 400.56 685 607 7.6 64.2 72.40 35184 9.70 10.40 5358 11.4 6237.00 91.2 90.10 49975 8.8 9.90 4798 3.4 3.80 1839 0.7 0.80 400 4238 560 560 30.3241 35.0467 39.7694 temperate deciduous malacophyll broadleaf forest 1166.45 401.20 833 749 7.3 68.2 75.80 44860 9.90 10.10 6556 10.1 6634.00 91.8 90.90 60417 8.2 9.10 5370 2.8 3.10 1817 0 0.00 0 3870 2290 1580 29.5995 36.7089 43.8182 temperate deciduous malacophyll broadleaf forest 1168.24 401.84 731 623 11 69.7 81.70 43734 10.40 10.90 6530 14.7 9279.00 96.3 95.70 60486 3.7 4.30 2320 1.6 1.90 1031 0 0.00 0 3866 687 4124 21.4378 28.6906 35.9434 temperate deciduous malacophyll broadleaf forest 1170.02 402.48 847 727 39.3 38.7 44.90 49908 40.10 46.60 51738 14.2 18314.00 97.5 97.10 126064 2.5 2.90 3205 0.4 0.40 458 0 0.00 0 5342 458 1831 26.642 29.9172 33.1925 cool mixed evergreen needleleaf and deciduous broadleaf forest 1171.80 403.12 900 786 34.9 35.7 41.20 34402 40.80 46.60 39101 12.7 12179.00 93.9 93.00 90276 6.1 7.00 5876 3.2 3.70 3099 0 0.00 0 7158 1496 1923 29.0591 33.3747 37.6903 cool mixed evergreen needleleaf and deciduous broadleaf forest 1174.79 403.76 923 740 34.2 31.3 39.10 27613 36.60 45.70 32294 19.8 17484.00 94.6 93.20 83409 5.4 6.80 4777 1.7 2.20 1529 0.1 0.10 96 4395 1815 1433 31.2409 35.3539 39.4669 cool mixed evergreen needleleaf and deciduous broadleaf forest 1176.38 404.08 849 685 34.7 34.6 42.90 37887 37.90 46.30 41494 19.4 21134.00 97.2 96.50 106310 2.8 3.50 3093 1 1.30 1160 0.5 0.60 515 3995 0 644 28.0853 32.3263 36.5672 cool mixed evergreen needleleaf and deciduous broadleaf forest 1177.96 404.40 805 693 35 42 48.90 19891 33.40 37.70 15782 13.9 6571.00 96.4 95.80 45532 3.6 4.20 1702 1.6 1.80 763 0.4 0.40 176 2171 1114 939 24.7417 33.4194 42.0971 cool evergreen needleleaf forest 1179.55 404.72 944 760 37.8 43.9 54.70 31939 28.40 34.10 20578 19.5 14128.00 98.4 98.00 71329 1.6 2.00 1152 0.3 0.40 230 0.4 0.50 307 3302 537 768 25.7832 29.3157 32.8481 temperate deciduous malacophyll broadleaf forest 1181.14 405.04 837 696 34.2 42.2 51.00 28001 26.60 31.20 17512 16.9 11122.00 95.5 94.50 63021 4.5 5.50 2997 2 2.40 1341 0.5 0.60 316 3234 2367 394 28.9949 34.2514 39.5078 temperate deciduous malacophyll broadleaf forest 1182.74 405.36 924 743 21.4 46 57.10 62191 18.00 21.80 24438 19.5 26487.00 96.3 95.40 130238 3.7 4.60 4975 0.5 0.60 731 0 0.00 0 4390 2780 1463 29.4578 34.686 39.9141 temperate deciduous malacophyll broadleaf forest 1184.32 405.68 747 570 13.2 57.9 76.20 30519 9.40 12.50 4993 23.7 12447.00 97.6 96.80 51266 2.4 3.20 1266 0.9 1.20 492 0.1 0.20 70 1195 1266 1055 15.05 25.1339 35.2178 temperate deciduous malacophyll broadleaf forest 1186.22 406.00 878 618 4 53.9 76.60 51891 14.10 16.90 13604 29.6 28524.00 97 95.80 93471 3 4.20 2852 0.2 0.30 219 0.1 0.20 110 4827 2524 1097 15.05 25.1339 35.2178 cool mixed evergreen needleleaf and deciduous broadleaf forest 1188.38 406.32 874 654 7 46.7 62.40 16765 24.70 30.80 8876 25.2 9040.00 95.5 94.00 34314 4.5 6.00 1603 1.3 1.70 452 0.1 0.20 41 2630 534 1192 21.5207 27.3245 33.1282 cool mixed evergreen needleleaf and deciduous broadleaf forest 1190.54 406.64 922 628 6.7 52.7 77.40 41263 9.00 10.60 7047 31.8 24960.00 93.4 90.30 73099 6.6 9.70 5179 0.6 1.00 510 0.4 0.60 340 4330 850 2462 27.1906 31.9252 36.6599 cool mixed evergreen needleleaf and deciduous broadleaf forest 1192.72 406.96 1137 462 1.3 11.9 28.60 2970 3.20 4.60 760 59.4 15072.00 74 35.90 18779 26 64.10 6610 2.9 7.10 737 1.8 4.30 469 1206 804 335 21.2539 27.7012 34.1484 temperate deciduous malacophyll broadleaf forest 1194.75 407.28 1113 522 3.1 34.4 73.00 86435 11.50 11.80 29039 53.1 134075.00 94.1 87.40 237524 5.9 12.60 14973 1.4 3.00 3630 0.1 0.20 227 7713 1134 2495 26.3939 28.5929 30.7919 cool mixed evergreen needleleaf and deciduous broadleaf forest 1196.67 407.60 1648 601 3.1 23 63.10 15926 8.80 10.70 6050 63.5 44000.00 91.4 76.40 63290 8.6 23.60 5968 2.3 6.10 1555 0.2 0.70 168 1555 925 252 20.4138 32.9574 45.501 cool evergreen needleleaf forest 1198.56 407.92 1281 481 3.5 9.8 26.00 6365 14.50 15.10 9370 62.5 40737.00 79.9 46.40 52092 20.1 53.60 13138 7.3 19.50 4787 0.2 0.40 102 7282 3819 6161 28.3848 33.2485 38.1123 cool evergreen needleleaf forest 1200.45 408.24 2600 516 2.4 6.7 34.30 3068 17.80 41.40 8024 80.1 36112.00 95.7 78.10 43095 4.3 21.90 1958 1.2 6.00 538 0.2 1.00 87 3518 347 260 17.0011 25.508 34.0148 cool evergreen needleleaf forest 1202.47 408.56 1354 506 5.4 14.2 38.20 3767 24.70 42.20 6496 62.7 16537.00 93.5 82.60 24688 6.5 17.40 1716 1 2.60 254 0.4 1.20 117 3023 20 449 14.3752 30.7504 47.1256 cool evergreen needleleaf forest 1204.66 408.88 1043 181 2.8 3.3 18.30 741 11.40 27.10 2652 82.6 19380.00 91.2 49.20 21382 8.8 50.80 2068 2.9 17.10 697 0 0.00 0 2923 315 292 20.9226 24.3628 27.803 cool evergreen needleleaf forest 1206.86 409.20 1865 518 2.4 12.3 43.90 4792 8.70 15.70 3421 72.2 28441.00 89.1 60.80 35092 10.9 39.20 4286 1.3 4.90 528 0.1 0.20 21 1668 0 253 20.4076 26.9838 33.5601 cool mixed evergreen needleleaf and deciduous broadleaf forest 1209.05 409.52 3287 480 0.9 6.8 46.70 2385 4.50 22.30 1505 85.4 29758.00 95.8 71.00 33373 4.2 29.00 1474 1.1 7.50 381 0.2 1.20 64 997 106 117 15.6783 20.6238 25.5692 cool mixed evergreen needleleaf and deciduous broadleaf forest 1212.09 410.16 596 181 7.1 7.1 23.20 1083 7.30 14.40 1109 69.7 10695.00 85.1 50.80 13066 14.9 49.20 2294 4.7 15.40 722 0.5 1.60 77 696 258 0 25.8964 29.1317 32.367 cool mixed evergreen needleleaf and deciduous broadleaf forest 1212.71 410.32 730 278 2.8 13.4 35.90 3967 14.20 16.90 4128 61.9 18120.00 82.9 55.00 24254 17.1 45.00 5011 1.8 4.70 521 1.5 3.80 441 1804 80 80 24.8396 36.9341 49.0286 cool evergreen needleleaf forest 1214.57 410.80 817 463 21.4 35.1 61.50 17257 16.70 21.70 8267 43.3 21360.00 92.2 86.20 45437 7.8 13.80 3862 1.7 3.00 845 1 1.70 483 1569 784 905 24.9655 36.7407 48.5159 temperate deciduous malacophyll broadleaf forest 1215.17 410.96 955 288 1.8 17.1 56.80 13988 11.60 11.00 9465 69.9 56882.00 91.2 70.80 74280 8.8 29.20 7164 1.7 5.50 1364 0.5 1.70 426 682 85 0 28.4339 33.511 38.5881 cool evergreen needleleaf forest 1217.00 411.44 855 129 3.1 3.2 21.00 459 15.00 32.00 2191 84.9 12325.00 93.2 55.00 13530 6.8 45.00 985 2.7 17.80 391 0.2 1.50 34 1188 204 51 18.7729 22.927 27.0812 cool evergreen needleleaf forest 1219.42 412.08 835 365 24.6 12.1 28.00 6963 29.80 59.60 17069 56.3 32088.00 95.2 89.00 54276 4.8 11.00 2731 1.4 3.20 820 0 0.00 0 6349 137 956 24.1935 28.1485 32.1034 cool evergreen needleleaf forest 1221.87 412.72 753 447 16.4 38 64.00 28221 19.10 29.30 14210 40.6 30195.00 97.5 95.70 72428 2.5 4.30 1875 1.1 1.70 789 1.1 1.80 789 2171 1184 987 26.4811 30.2271 33.9731 cool mixed evergreen needleleaf and deciduous broadleaf forest 1224.34 413.36 742 586 28.1 41.8 53.00 28373 31.40 39.60 21415 21.1 14277.00 96 94.90 65161 4 5.10 2746 1.2 1.60 824 0 0.00 0 1190 1190 92 22.4264 32.7333 43.0401 cool mixed evergreen needleleaf and deciduous broadleaf forest 1226.69 414.00 779 702 33.9 47.5 52.60 35014 38.70 43.00 28657 9.9 7307.00 97.7 97.40 72213 2.3 2.60 1708 0.9 1.00 664 0 0.00 0 1803 0 95 23.8546 27.487 31.1194 cool mixed evergreen needleleaf and deciduous broadleaf forest 1229.03 414.64 830 737 38 47.8 53.80 46465 37.30 42.00 36283 11.2 10885.00 98.3 98.10 95506 1.7 1.90 1639 0.4 0.40 351 0 0.00 0 5033 234 234 20.9368 27.7382 34.5396 cool mixed evergreen needleleaf and deciduous broadleaf forest 1231.06 415.28 854 742 48 40.1 45.80 61785 43.10 49.40 66675 13.1 20292.00 98.6 98.40 152557 1.4 1.60 2174 0 0.00 0 0 0.00 0 7972 1087 1268 24.8442 28.5884 32.3325 cool mixed evergreen needleleaf and deciduous broadleaf forest 1232.94 415.91 703 561 41 36.6 46.10 44328 40.10 50.60 48623 20.2 24397.00 98 97.50 118376 2 2.50 2405 0.4 0.60 516 1.1 1.40 1374 4811 0 172 14.3502 28.0403 41.7304 cool mixed evergreen needleleaf and deciduous broadleaf forest 1234.72 416.56 704 598 47 34.3 40.40 32238 47.70 56.00 44629 15.1 14121.00 98.6 98.30 92455 1.4 1.70 1332 0.6 0.70 533 0 0.00 0 2798 133 533 17.046 23.188 29.3299 cool mixed evergreen needleleaf and deciduous broadleaf forest 1236.62 417.20 758 616 35.5 40.6 49.90 27075 37.20 45.50 24871 18.8 12523.00 97.9 97.40 65441 2.1 2.60 1411 0.8 1.00 529 0.1 0.20 88 2469 0 176 24.1138 27.0237 29.9335 cool mixed evergreen needleleaf and deciduous broadleaf forest 1238.50 417.84 800 648 11.5 59.8 73.90 46060 15.30 18.90 11731 19 14616.00 96.3 95.40 74044 3.8 4.60 2885 0.9 1.10 673 0 0.00 0 1923 1442 192 20.2559 23.2379 26.2198 cool mixed evergreen needleleaf and deciduous broadleaf forest 1240.38 418.48 704 573 14.8 56.8 69.70 43739 21.30 25.80 16402 18.6 14325.00 97.7 97.20 75231 2.3 2.80 1750 0.9 1.00 656 0.7 0.90 547 1859 1093 437 13.7534 21.6872 29.6209 cool mixed evergreen needleleaf and deciduous broadleaf forest 1242.31 419.12 683 519 8.6 56.3 73.80 15473 11.90 15.30 3265 24 6609.00 93.4 91.30 25708 6.6 8.70 1813 3.2 4.30 887 0.6 0.80 161 3425 40 685 22.8806 29.9078 36.935 temperate deciduous malacophyll broadleaf forest 1244.30 419.76 870 302 0.8 18.1 52.40 3823 4.00 4.00 847 65.2 13746.00 86.1 59.90 18126 13.9 40.10 2928 4.1 11.50 847 0.9 2.60 194 1234 48 3291 18.5374 23.1219 27.7064 cool mixed evergreen needleleaf and deciduous broadleaf forest 1246.14 420.40 868 253 1.4 10.7 37.10 1461 2.70 5.20 374 70.9 9547.00 84.3 46.20 11363 15.7 53.80 2111 6.8 23.30 916 0.7 2.30 93 1040 78 264 18.2578 24.8153 31.3729 graminoids with forbs 1247.87 421.04 3569 409 1.1 4.7 39.80 1072 10.40 24.70 2446 88.5 20779.00 96.4 68.20 22613 3.6 31.80 855 1 8.80 237 0.1 0.50 13 460 46 112 11.3568 19.2923 27.2278 cool evergreen needleleaf forest 1250.02 421.68 2727 240 2 0.5 7.50 102 6.80 41.20 1055 91.2 13937.00 95.8 52.10 14637 4.2 47.90 644 1.5 17.10 230 0 0.00 0 644 11 22 9.1801 15.1719 21.1637 cool evergreen needleleaf forest 1252.82 422.32 2497 328 1 2.7 20.70 958 8.00 45.10 2834 86.8 30586.00 95.8 68.30 33745 4.2 31.70 1467 1.7 13.50 620 0 0.30 14 2637 1720 197 16.1665 18.4912 20.816 cool evergreen needleleaf forest 1254.19 422.64 1391 328 3.4 6.1 25.60 3018 14.70 36.90 7324 76.5 38166.00 91.7 64.90 45813 8.3 35.10 4129 1.8 7.60 897 0.3 1.20 144 7432 0 467 14.8431 26.6813 38.5196 cool evergreen needleleaf forest 1255.89 422.96 2769 222 0.3 3.2 41.30 1164 1.60 18.60 575 91.9 32575.00 96.9 60.80 34301 3.1 39.20 1113 0.8 9.10 256 0.9 9.80 307 486 64 38 9.8964 15.8633 21.8302 cool mixed evergreen needleleaf and deciduous broadleaf forest 1257.70 423.28 1562 624 1.6 10.8 27.00 1634 6.50 4.70 993 60.1 9135.00 74.2 35.40 11288 25.8 64.60 3925 2.5 6.20 379 0.6 1.40 88 682 0 58 19.0053 30.8622 42.7192 graminoids with forbs 1259.52 423.60 1629 611 2.3 23.8 63.60 3455 3.70 8.20 551 62.5 9038.00 90.4 74.50 13078 9.6 25.50 1385 2.6 6.70 364 0.3 0.80 44 311 204 80 18.3447 32.5955 46.8464 temperate deciduous malacophyll broadleaf forest 1261.32 423.92 616 450 2.2 13.7 18.70 2439 6.10 3.80 1103 26.9 4822.00 44 23.30 7873 56 76.70 10022 4.4 6.00 784 1 1.30 174 1046 0 0 24.8667 27.8202 30.7737 graminoids with forbs 1262.94 424.24 1510 366 2.1 13.9 56.40 4926 8.40 27.60 3060 75.8 27349.00 96.3 84.70 34761 3.7 15.30 1339 1.6 6.60 574 0.5 1.90 167 1626 24 0 14.4082 25.326 36.2438 temperate deciduous malacophyll broadleaf forest 1264.51 424.56 745 495 1.5 22.2 33.50 10073 5.70 3.20 2610 33.5 15172.00 59.1 38.40 26702 40.9 61.60 18509 2.4 3.60 1092 0.7 1.00 303 3520 61 425 21.9924 32.4287 42.8649 cool mixed evergreen needleleaf and deciduous broadleaf forest 1266.10 424.88 722 430 9.3 27.3 46.00 14477 29.70 47.10 15646 40.4 21351.00 97 94.90 51184 3 5.10 1609 0.4 0.70 219 0.3 0.50 146 1974 0 4095 18.0026 31.7524 45.5022 cool evergreen needleleaf forest 1267.69 425.20 891 552 2.8 28.5 46.10 34102 8.20 8.70 9800 38 45513.00 74.2 58.30 88744 25.8 41.70 30879 5.6 9.00 6713 0.2 0.40 269 10606 0 269 24.6087 30.4461 36.2835 cool mixed evergreen needleleaf and deciduous broadleaf forest 1270.58 425.52 702 253 10.1 15.4 43.20 10668 21.10 45.80 14485 64 43942.00 96.3 89.70 66157 3.7 10.30 2545 0.9 2.40 587 0.6 1.60 391 3621 587 685 18.4672 26.4517 34.4362 cool evergreen needleleaf forest 1272.39 425.68 1175 657 10.3 31.5 56.30 34080 19.80 31.50 21370 44.1 47711.00 93.9 89.00 101593 6.1 11.00 6632 2.5 4.60 2763 0.2 0.30 184 6908 92 2579 27.8077 29.0822 30.3566 cool mixed evergreen needleleaf and deciduous broadleaf forest 1274.22 425.84 1109 524 11.7 27.6 58.30 35138 19.70 35.00 25230 52.8 67399.00 98.2 96.20 125466 1.8 3.80 2304 1.1 2.30 1382 0.1 0.20 115 6221 2765 2650 21.1759 25.2337 29.2915 cool mixed evergreen needleleaf and deciduous broadleaf forest 1276.05 426.00 935 604 15.8 40.3 62.50 98687 20.00 28.00 49084 35.4 86418.00 96.7 94.90 236016 3.3 5.10 8093 1.3 2.00 3133 0.1 0.20 261 6266 2872 8877 25.6551 29.8355 34.0159 cool mixed evergreen needleleaf and deciduous broadleaf forest 1277.88 426.16 720 403 23.2 18 32.20 42978 34.80 58.20 82650 44 104800.00 97.8 96.00 232742 2.2 4.00 5290 1.1 1.90 2645 0.8 1.50 1984 12232 8265 8265 14.4498 22.2777 30.1057 cool evergreen needleleaf forest 1279.71 426.32 1101 646 18.7 23.4 40.00 69258 30.60 47.90 90197 41.4 122143.00 97.4 95.50 287776 2.6 4.50 7785 0.6 1.00 1610 0.2 0.30 537 13422 10469 11812 29.7954 30.8646 31.9339 cool mixed evergreen needleleaf and deciduous broadleaf forest 1281.54 426.48 1092 710 21.5 27.4 42.00 35186 33.20 44.80 42715 35 44952.00 96.3 94.40 123793 3.7 5.60 4707 1.5 2.20 1883 0.5 0.70 588 5766 5060 4001 27.4045 29.3591 31.3136 cool mixed evergreen needleleaf and deciduous broadleaf forest 1284.19 426.80 735 473 22 26.8 41.80 17072 35.00 53.40 22247 35.7 22592.00 98 96.80 62086 2 3.20 1293 0.6 1.00 431 0.5 0.80 345 1035 5777 3018 14.8723 25.2651 35.6579 cool evergreen needleleaf forest 1286.12 427.12 1088 813 16.5 36.5 48.50 33448 28.50 35.70 26165 25.3 23286.00 90.1 86.70 82985 9.9 13.30 9145 2.2 3.00 2032 0 0.00 0 7028 2710 2371 27.5642 30.819 34.0739 cool mixed evergreen needleleaf and deciduous broadleaf forest 1288.08 427.44 809 375 7.7 18.8 41.20 15809 24.40 51.70 20324 53.7 44548.00 97.2 93.90 80679 2.8 6.10 2361 0.8 1.80 718 0.1 0.30 103 4619 11702 6261 17.643 26.4311 35.2192 cool evergreen needleleaf forest 1289.03 427.60 1031 414 4.3 22.1 54.70 24902 14.50 24.70 16346 59.9 67683.00 93.2 83.10 105418 6.8 16.90 7679 1.2 2.90 1317 0.2 0.50 219 2194 19306 1426 27.4442 31.5402 35.6362 cool evergreen needleleaf forest 1291.54 428.08 1106 307 4 17.4 62.80 13901 10.40 24.40 8325 72.2 57841.00 96.7 88.30 77458 3.3 11.70 2606 1.6 5.50 1231 1.2 4.10 941 3113 4923 3402 21.635 23.8834 26.1319 cool evergreen needleleaf forest 1293.80 428.72 797 378 2.8 11.5 24.10 1485 6.10 10.30 783 52.5 6840.00 69.8 36.20 9076 30.2 63.80 3934 11.8 24.80 1534 0.1 0.30 16 82 3787 1436 27.5167 31.0865 34.6562 cool mixed evergreen needleleaf and deciduous broadleaf forest 1296.08 429.36 957 412 7.9 15.9 37.00 7595 11.60 24.90 5647 56.9 27229.00 85.5 66.30 40868 14.5 33.70 6945 8.3 19.40 3997 1.4 3.30 699 3647 13489 4147 29.3604 33.7418 38.1231 cool evergreen needleleaf forest 1298.48 430.00 1207 432 2.8 8.8 24.50 1263 10.40 10.80 1476 64.2 9224.00 77.4 36.80 11116 22.6 63.20 3249 6.6 18.20 940 1 2.70 143 357 655 202 21.7995 28.5312 35.263 graminoids with forbs 1300.91 430.64 1013 488 4 15.2 31.40 1698 5.80 9.10 638 51.8 5792.00 72.9 43.60 8142 27.1 56.40 3034 10.3 21.30 1148 1.6 3.20 177 154 2361 1666 34.2203 39.1789 44.1376 cool mixed evergreen needleleaf and deciduous broadleaf forest 1303.62 431.28 735 554 7.5 47 62.20 19655 20.70 26.20 8661 24.7 10311.00 92.8 90.40 38852 7.2 9.60 3019 2.6 3.40 1082 0.7 0.90 285 1139 11906 8146 26.6541 35.9394 45.2248 cool mixed evergreen needleleaf and deciduous broadleaf forest 1306.18 431.92 1378 445 1.6 15.9 48.70 3601 4.70 8.10 1046 67.7 15478.00 86.5 58.20 19775 13.5 41.80 3086 6.2 19.10 1410 0.1 0.40 33 1228 25017 2372 13.3898 26.6269 39.8641 cool mixed evergreen needleleaf and deciduous broadleaf forest 1308.28 432.56 921 570 9.4 32.3 52.30 19559 22.30 35.30 13567 38.1 23114.00 95.4 92.60 57885 4.6 7.40 2766 1.6 2.70 988 1.4 2.20 856 1251 263 1054 21.3227 24.6352 27.9477 cool mixed evergreen needleleaf and deciduous broadleaf forest 1310.38 433.20 1298 861 13.2 33.3 49.90 18333 25.00 36.70 13814 33.6 18631.00 94.6 91.90 52355 5.4 8.10 2984 3.3 5.00 1834 3.8 5.60 2174 3965 43 1023 27.0936 33.4594 39.8252 cool mixed evergreen needleleaf and deciduous broadleaf forest 1312.60 433.84 939 530 13.5 24.5 43.70 12079 26.70 46.00 13125 43.5 21387.00 96.2 93.20 47219 3.8 6.80 1882 2.6 4.70 1308 1.4 2.40 680 1935 52 732 20.8991 24.7181 28.5371 cool mixed evergreen needleleaf and deciduous broadleaf forest 1314.81 434.48 908 646 14.7 33.9 48.00 20226 29.30 39.60 17411 28.8 17151.00 95 93.00 56493 5 7.00 2946 2.1 3.00 1243 1.2 1.70 720 2226 720 1506 24.4176 28.329 32.2404 cool mixed evergreen needleleaf and deciduous broadleaf forest 1317.11 435.12 800 504 17.5 24.6 38.90 15621 29.90 46.70 19048 37 23590.00 96.8 94.80 61685 3.3 5.20 2072 2 3.20 1275 1.1 1.80 717 1355 558 4702 26.6777 30.2386 33.7996 cool mixed evergreen needleleaf and deciduous broadleaf forest 1319.40 435.76 924 586 10 22.7 35.80 16665 34.30 49.90 25235 36.6 26822.00 93.3 89.40 68403 6.7 10.60 4920 2 3.30 1507 0.1 0.20 79 2777 1270 1904 29.739 34.1317 38.5244 cool evergreen needleleaf forest 1321.54 436.40 899 377 7.4 16.5 38.90 7692 26.10 36.50 12240 58.1 27304.00 91.4 79.60 42997 8.6 20.40 4028 4.5 10.80 2145 0.4 1.00 209 1726 2145 2354 28.0961 33.6793 39.2625 cool evergreen needleleaf forest 1323.92 437.04 598 369 2 15.8 25.50 2152 7.40 5.20 1007 38.3 5239.00 58.9 33.30 8053 41.1 66.70 5628 13.5 21.90 1853 2.4 3.90 343 572 389 69 31.9527 36.1501 40.3474 cool mixed evergreen needleleaf and deciduous broadleaf forest 1325.84 437.20 706 502 3.20 28.70 40.50 1605.00 7.50 7.40 426.00 28.90 1612.00 64.30 49.80 3589.00 35.70 50.20 1992.00 14.60 20.50 814.00 3.70 5.10 213.00 150.00 8.00 40.00 27.6287 38.6196 49.6105 temperate deciduous malacophyll broadleaf forest 1327.75 437.36 69 42 2.80 11.50 19.10 378.00 10.10 16.70 331.00 39.10 1277.00 65.20 42.90 2128.00 34.80 57.10 1135.00 8.70 14.30 284.00 8.00 12.50 284.00 9645.00 0.00 0.00 -999 -999 -999 graminoids with forbs 1329.67 437.52 737 372 7.30 10.30 20.70 919.00 13.20 18.20 1155.00 49.50 4345.00 73.40 47.30 6441.00 26.60 52.70 2334.00 9.00 17.70 786.00 1.20 2.40 107.00 4536.00 12.00 0.00 29.2543 45.6956 62.1369 cool evergreen needleleaf forest 1332.20 437.68 477 240 9.80 17.30 34.60 924.00 17.40 18.70 925.00 49.70 2645.00 78.60 57.50 4186.00 21.40 42.50 1139.00 7.60 15.00 402.00 2.70 5.10 145.00 1250.00 100.00 11.00 33.9929 36.3164 38.64 cool evergreen needleleaf forest 1336.05 438.00 203 97 7.40 17.20 36.10 1265.00 16.30 22.60 1195.00 52.20 3836.00 82.30 62.90 6044.00 17.70 37.10 1303.00 7.40 15.40 543.00 2.40 4.90 181.00 3728.00 0.00 36.00 -999 -999 -999 cool evergreen needleleaf forest 1337.96 438.16 401 224 6.60 12.70 23.00 959.00 19.40 17.80 1441.00 44.20 3271.00 70.30 46.90 5211.00 29.70 53.10 2199.00 10.70 19.20 795.00 1.50 2.60 111.00 2606.00 0.00 18.00 45 45 45 cool evergreen needleleaf forest 1339.46 438.32 290 139 4.00 12.20 25.90 770.00 13.70 19.40 855.00 52.10 3233.00 75.20 48.20 4668.00 24.80 51.80 1542.00 13.50 28.00 835.00 2.00 4.10 128.00 450.00 471.00 43.00 -999 -999 -999 cool evergreen needleleaf forest 1341.94 438.96 351 169 10.30 12.80 26.80 1787.00 15.10 27.90 2105.00 51.80 7232.00 79.80 58.00 11127.00 20.20 42.00 2821.00 11.90 24.90 1669.00 3.00 6.10 437.00 1510.00 596.00 40.00 -999 -999 -999 cool evergreen needleleaf forest 1344.67 439.60 739 606 23.40 32.50 39.80 12123.00 11.00 12.60 4125.00 18.00 6691.00 88.40 85.80 32849.00 11.60 14.20 4326.00 6.30 7.80 2364.00 1.30 1.60 503.00 1610.00 2716.00 1308.00 27.9043 41.1849 54.4656 temperate deciduous malacophyll broadleaf forest 1345.14 439.76 771 298 1.10 10.30 26.50 5067.00 16.00 10.40 7888.00 61.30 30335.00 78.90 45.30 38994.00 21.10 54.70 10454.00 10.90 28.20 5387.00 0.80 2.00 385.00 1796.00 9428.00 5131.00 25.0788 27.4554 29.8321 cool evergreen needleleaf forest 1346.45 440.24 798 668 26.80 43.40 51.60 17421.00 23.20 24.60 9316.00 16.30 6546.00 93.40 92.10 37511.00 6.60 7.90 2669.00 4.20 4.90 1661.00 2.20 2.60 906.00 302.00 554.00 403.00 32.3244 35.6677 39.011 temperate deciduous malacophyll broadleaf forest 1348.20 440.88 767 686 18.70 53.10 59.30 20165.00 15.70 16.00 5981.00 10.50 4004.00 91.80 90.80 34799.00 8.20 9.20 3114.00 3.40 3.70 1285.00 1.50 1.70 593.00 544.00 99.00 99.00 36.5108 40.4018 44.2927 temperate deciduous malacophyll broadleaf forest 1349.92 441.52 744 678 14.30 63.30 69.30 135368.00 14.30 15.70 30752.00 8.90 18969.00 91.10 90.30 194863.00 8.90 9.70 18969.00 4.20 4.50 8909.00 0.50 0.60 1150.00 2299.00 33052.00 575.00 25.7354 38.9576 52.1798 temperate deciduous malacophyll broadleaf forest 1351.65 442.16 890 786 17.30 53.90 60.90 12287.00 17.60 20.00 4020.00 11.70 2662.00 90.00 88.70 20502.00 10.00 11.30 2278.00 6.50 7.40 1484.00 0.60 0.60 128.00 307.00 793.00 230.00 28.8327 34.3408 39.849 temperate deciduous malacophyll broadleaf forest 1353.25 442.80 735 699 11.20 62.50 65.70 17247.00 12.50 13.00 3459.00 4.90 1353.00 86.30 85.60 23823.00 13.70 14.40 3795.00 6.20 6.60 1729.00 1.20 1.30 338.00 451.00 977.00 75.00 24.352 39.6049 54.8578 cool mixed evergreen needleleaf and deciduous broadleaf forest 1354.81 443.44 737 707 11.70 63.30 66.00 14348.00 11.60 12.10 2642.00 4.10 922.00 86.20 85.60 19509.00 13.80 14.40 3134.00 5.90 6.20 1352.00 1.70 1.80 399.00 184.00 3257.00 31.00 25.4337 38.4228 51.4119 cool mixed evergreen needleleaf and deciduous broadleaf forest 1356.53 444.08 757 712 10.80 56.60 60.10 11898.00 12.40 12.40 2614.00 6.00 1251.00 82.70 81.60 17402.00 17.30 18.40 3642.00 8.20 8.70 1723.00 1.20 1.20 250.00 334.00 9118.00 139.00 37.4991 41.0866 44.6741 temperate deciduous malacophyll broadleaf forest 1358.28 444.72 753 709 12.10 59.70 63.70 72812.00 16.40 17.40 20018.00 5.80 7104.00 87.30 86.50 106071.00 12.70 13.50 15499.00 5.90 6.20 7104.00 0.90 1.00 1130.00 1776.00 118018.00 807.00 31.3356 35.5404 39.7451 cool mixed evergreen needleleaf and deciduous broadleaf forest 1359.93 445.36 754 733 7.70 65.20 67.10 19149.00 8.30 8.50 2452.00 2.80 817.00 81.70 81.20 23973.00 18.30 18.80 5371.00 8.90 9.20 2608.00 1.80 1.90 545.00 311.00 17746.00 234.00 33.1994 37.1606 41.1218 cool mixed evergreen needleleaf and deciduous broadleaf forest 1361.61 446.00 617 586 9.20 62.20 65.40 16735.00 12.70 12.30 3399.00 5.10 1351.00 83.80 82.90 22532.00 16.20 17.10 4358.00 5.60 6.00 1525.00 1.80 1.80 479.00 1395.00 1525.00 305.00 35.3968 38.7262 42.0556 cool mixed evergreen needleleaf and deciduous broadleaf forest 1363.18 446.64 761 694 12.00 54.00 59.20 15930.00 15.70 17.30 4652.00 8.80 2597.00 89.60 88.60 26437.00 10.40 11.40 3062.00 4.70 5.20 1395.00 1.30 1.40 388.00 388.00 78.00 930.00 34.0597 37.4636 40.8675 cool mixed evergreen needleleaf and deciduous broadleaf forest 1365.00 447.28 1303 838 18.70 19.50 29.90 46834.00 22.80 28.60 55197.00 35.70 86419.00 77.70 65.30 188079.00 22.30 34.70 54082.00 3.80 6.00 9293.00 7.10 10.60 18399.00 40143.00 0.00 1858.00 40.1103 44.4219 48.7335 cool evergreen needleleaf forest