# Great Salt Lake, Utah Biomarker Data from the Holocene #----------------------------------------------------------------------- # World Data Service for Paleoclimatology, Boulder # and # NOAA Paleoclimatology Program #----------------------------------------------------------------------- # Template Version 4.0 # Encoding: UTF-8 # NOTE: Please cite original publication, NOAA Landing Page URL, dataset and publication DOIs (where available), and date accessed when using downloaded data. # If there is no publication information, please cite investigator, study title, NOAA Landing Page URL, and date accessed. # # Description/Documentation lines begin with '#' followed by a space # Data lines have no '#' # # NOAA_Landing_Page: https://www.ncei.noaa.gov/access/paleo-search/study/36833 # Landing_Page_Description: NOAA Landing Page of this file's parent study, which includes all study metadata. # # Study_Level_JSON_Metadata: https://www.ncei.noaa.gov/pub/data/metadata/published/paleo/json/noaa-lake-36833.json # Study_Level_JSON_Description: JSON metadata of this data file's parent study, which includes all study metadata. # # Data_Type: Paleolimnology # # Dataset_DOI: 10.25921/f3hg-4e71 # # Science_Keywords: Air Temperature Reconstruction #--------------------------------------- # Resource_Links # # Data_Download_Resource: https://www.ncei.noaa.gov/pub/data/paleo/paleolimnology/northamerica/usa/utah/so2022/so2022-glad1-gsl00-1b_biomarker.txt # Data_Download_Description: NOAA Template File; Biomarker Data # #--------------------------------------- # Contribution_Date # Date: 2022-09-12 #--------------------------------------- # File_Last_Modified_Date # Date: 2024-05-13 #--------------------------------------- # Title # Study_Name: Great Salt Lake, Utah Biomarker Data from the Holocene #--------------------------------------- # Investigators # Investigators: So, Rachel; Lowenstein, Tim; Jagniecki, Elliot; Tierney, Jessica; Feakins, Sarah #--------------------------------------- # Description_Notes_and_Keywords # Description: # Provided Keywords: biomarker, GDGT #--------------------------------------- # Publication # Authors: Rachel T. So, Tim K. Lowenstein, Elliot Jagniecki, Jessica E. Tierney, Sarah J. Feakins # Published_Date_or_Year: 2023-06-01 # Published_Title: Holocene water balance variations in Great Salt Lake, Utah: application of GDGT indices and the ACE salinity proxy # Journal_Name: Paleoceanography and Paleoclimatology # Volume: 38 # Edition: # Issue: 6 # Pages: # Report_Number: e2022PA004558 # DOI: 10.1029/2022PA004558 # Online_Resource: # Full_Citation: # Abstract: Great Salt Lake, Utah, is a hypersaline terminal lake in the Great Basin, and the remnant of the late glacial Lake Bonneville. Holocene hydroclimate variations cannot be interpreted from the shoreline record, but instead can be investigated by proxies archived in the sediments. GLAD1-GSL00-1B was cored in 2000 and recently dated by radiocarbon for the Holocene section with the top 11 m representing ~7 ka to present. Sediment samples every 30 cm (~220 years) were studied for the full suite of microbial membrane lipids, including those responsive to temperature and salinity. The Archaeol and Caldarchaeol Ecometric (ACE) index detects the increase in lipids of halophilic archaea, relative to generalists, as salinity increases. We find Holocene ACE values ranged from 81-98, which suggests persistent hypersalinity with <50 g/L variability across 7.2 ka. The temperature proxy, MBTʹ5Me,­ yields values similar to modern mean annual air temperature for months above freezing (MAF = 15.7°C) over the last 5.5 ka. Several GDGT metrics show a step shift in microbial communities and limnology at 5.5 ka. Extended archaeol detects elevated salinity during the regional mid-Holocene drought, not readily detected in the Holocene ACE record that is often near the upper limit of the salinity proxy. We infer that the mid-Holocene GSL was shallower and saltier than the late Holocene. The current drying may be returning the lake to conditions not seen since the mid-Holocene. #--------------------------------------- # Funding_Agency # Funding_Agency_Name: National Science Foundation # Grant: 1903665 #--------------------------------------- # Funding_Agency # Funding_Agency_Name: David and Lucile Packard Foundation # Grant: #--------------------------------------- # Site_Information # Site_Name: GLAD1-GSL00-1B (Great Salt Lake) # Location: Utah # Northernmost_Latitude: 41.094034 # Southernmost_Latitude: 41.094034 # Easternmost_Longitude: -112.364796 # Westernmost_Longitude: -112.364796 # Elevation_m: 1270 #--------------------------------------- # Data_Collection # Collection_Name: GLAD1-GSL00-1B_Biomarker # First_Year: 7206 # Last_Year: 291 # Time_Unit: calendar year before present # Core_Length_m: 33.13 # Parameter_Keywords: geochemistry, age control, lake level, physical properties # Notes: We report data for the middle to late Holocene section (upper 10.66 m) #--------------------------------------- # Chronology_Information # Chronology: #--------------------------------------- # Variables # PaST_Thesaurus_Download_Resource: https://www.ncei.noaa.gov/access/paleo-search/skos/past-thesaurus.rdf # PaST_Thesaurus_Download_Description: Paleoenvironmental Standard Terms (PaST) Thesaurus terms, definitions, and relationships in SKOS format. # # Data variables follow that are preceded by "##" in columns one and two. # Variables list, one per line, shortname-tab-var components: what, material, error, units, seasonality, data type, detail, method, C or N for Character or Numeric data) # ## depth_m depth,,,meter,,Paleolimnology; Climate Reconstructions,,,N,spliced depth ## age_calyrsBP age,,,calendar year before present,,Paleolimnology; Climate Reconstructions,,,N,Mean; central estimate of the age model. Age model derived using BACON based on radiocarbon chronology using all aquatic organic matter tie points (algal + cysts + short chain alkanes + TOC) of Bowen et al. (2019) Multi-substrate radiocarbon data constrain detrital and reservoir effects in Holocene sediments of the Great Salt Lake Utah. Radiocarbon 61: 905-926 (doi:10.1017/RDC.2019.62) ## ID sample identification,,,,,Paleolimnology,,,C,Core ID given by the Continental Scientific Drilling Facility; Core year hole section top depth-bottom depth ## depth_top depth,,,centimeter,,Paleolimnology,,,N, ## depth_bottom depth,,,centimeter,,Paleolimnology,,,N, ## archaeol archaeol,,,parts per billion,,Paleolimnology,,mass spectrometry,N, ## isoGDGT-0 isoprenoid glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of GDGT-0 ## isoGDGT-1 isoprenoid glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of GDGT-1 ## isoGDGT-2 isoprenoid glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of GDGT-2 ## isoGDGT-3 isoprenoid glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of GDGT-3 ## isoGDGT-5 isoprenoid glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of crenarchaeol ## isoGDGT-5_prime isoprenoid glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of crenarchaeol_prime ## b-IIIa branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIIa ## b-IIIa_prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIIa_prime; b-IIIa prime ## b-IIIa_3prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIIa_prime; b-IIIa 3 prime ## b-IIIb branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIIb ## b-IIIb_prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIIb_prime ## b-IIIc branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIIc ## bIIIc_prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIIc_prime ## b-IIa branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIa ## b-IIa_prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIa_prime; b-IIa prime ## b-IIa_3prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIa_prime; b-IIa 3 prime ## b-IIb branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIb ## b-IIb_prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIb_prime ## b-IIc branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIc ## b-IIc_prime branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-IIc_prime ## b-Ia branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-Ia ## b-Ib branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-Ib ## b-Ic branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of brGDGT-Ic ## isoGDGTs_ngg isoprenoid glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of summed isoprenoid GDGTs ## brGDGTs_ngg branched glycerol dialkyl glycerol tetraether,,,parts per billion,,Paleolimnology,,mass spectrometry,N,concentration of summed branched GDGTs ## R_ib organic compound index,,,dimensionless,,Paleolimnology,,mass spectrometry,N,ratio of the sum of brGDGTs to the sum of brGDGT and isoGDGT: R_i/b = sum(brGDGT)/(sum(isoGDGT)+sum(brGDGT)) ## IIIa_IIa organic compound index,,,dimensionless,,Paleolimnology,,mass spectrometry,N,index expressing relative proportion of brGDGT IIIa to brGDGT IIa: IIIa/IIa = (IIIa+IIIa')/(IIa+IIa'); Xiao et al. (2016) Ubiquitous production of branched glycerol dialkyl glycerol tetraethers (brGDGTs) in global marine environments: a new source indicator for brGDGTs. Biogeosciences 13: 5883-5894 (doi: 10.5194/bg-13-5883-2016) ## IIIa_IIa_7Me organic compound index,,,dimensionless,,Paleolimnology,,mass spectrometry,N,index expressing relative proportion of brGDGT IIIa to brGDGT Iia including 7 methyl isomers: IIIa/IIa = (IIIa+IIIa_prime+IIIa_3prime)/(IIa+IIa_prime+Iia_3prime); Martin et al. (2020) Early Holocene Thermal Maximum recorded by branched tetraethers and pollen in Western Europe (Massif Central France). Quaternary Science Reviews 228 (doi:10.1016/j.quascirev.2019.106109) ## BIT branched and isoprenoid tetraether index,,,dimensionless,,Paleolimnology,,mass spectrometry,N,BIT index; Hopmans et al. (2004) A novel proxy for terrestrial organic matter in sediments based on branched and isoprenoid tetraether lipids. Earth and Planetary Science Letters 224: 107-116 (doi:10.1016/j.epsl.2004.05.012) ## MBT5Me methylation index of branched tetraethers,,,dimensionless,,Paleolimnology,,mass spectrometry,N,pentamethylated brGDGT index MBTprime5Me; brGDGT index: MBT_prime_5Me = (Ia + Ib + Ic) / (Ia + Ib + Ic + IIa + IIb + IIc + IIIa); De Jonge et al.(2014) Occurrence and abundance of 6-methyl branched glycerol dialkyl glycerol tetraethers in soils: Implications for palaeoclimate reconstruction. Geochimica Et Cosmochimica Acta 141: 97-112 (doi:10.1016/j.gca.2014.06.013) ## MAT air temperature,glycerol dialkyl glycerol tetraether index,,degree Celsius,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,median; months above freezing mean annual air temperature using BayMBT0; Crampton-Flood et al. (2020) BayMBT: A Bayesian calibration model for branched glycerol dialkyl glycerol tetraethers in soils and peats. Geochimica Et Cosmochimica Acta 268: 142-159 (doi:10.1016/j.gca.2019.09.043) ## MAT_2.5th air temperature,glycerol dialkyl glycerol tetraether index,percentile,degree Celsius,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,2.5th percentile; months above freezing; mean annual air temperature; using BayMBT0 ## MAT_97.5th air temperature,glycerol dialkyl glycerol tetraether index,percentile,degree Celsius,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,97.5th percentile; months above freezing; mean annual air temperature; using BayMBT0 ## MATsoil air temperature,glycerol dialkyl glycerol tetraether index,,degree Celsius,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,median; months above freezing; mean annual air temperature; using BayMBT0; Crampton-Flood et al. (2020) BayMBT: A Bayesian calibration model for branched glycerol dialkyl glycerol tetraethers in soils and peats. Geochimica Et Cosmochimica Acta 268: 142-159 (doi:10.1016/j.gca.2019.09.043) ## MATsoil_2.5th air temperature,glycerol dialkyl glycerol tetraether index,percentile,degree Celsius,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,2.5th percentile; months above freezing; mean annual air temperature; using BayMBT0; soil calibration applied to lake sediments ## MATsoil_97.5th air temperature,glycerol dialkyl glycerol tetraether index,percentile,degree Celsius,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,97.5th percentile; months above freezing; mean annual air temperature; using BayMBT0; soil calibration applied to lake sediments ## CBTprime cyclization index of branched tetraethers,,,dimensionless,,Paleolimnology,,mass spectrometry,N,cyclization of hexamethyl branched tetraethers index; CBTprime; De Jonge et al. (2014) Occurrence and abundance of 6-methyl branched glycerol dialkyl glycerol tetraethers in soils: Implications for palaeoclimate reconstruction. Geochimica Et Cosmochimica Acta 141: 97-112 (doi:10.1016/j.gca.2014.06.013) ## pH pH,glycerol dialkyl glycerol tetraether index,,dimensionless,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,pH inferred using the linear regression of the lake calibration dataset (CBT_prime = 0.23(pH)-1.98) of pH and CBT_prime data reported in reference; Raberg et al. (2022) Near-universal trends in brGDGT lipid distributions in nature. Science Advances 8:12 (doi: 10.1126/sciadv.abm7625) ## pHsoil pH,glycerol dialkyl glycerol tetraether index,,dimensionless,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,pH inferred using the linear regression of the soil calibration dataset (CBT_prime = 0.50(pH)-3.65) of pH and CBT_prime data reported in reference; Raberg et al. (2022) Near-universal trends in brGDGT lipid distributions in nature. Science Advances 8: 12 (doi: 10.1126/sciadv.abm7625) ## ACE archaeol caldarchaeol ecometric,,,percent,,Paleolimnology,,mass spectrometry,N,Archeol Caldarcheol Ecometric; ACE = [archaeol]/[archaeol + caldarchaeol] x 100 where caldarchaeol = isoGDGT-0. Turich and Freeman (2011). Archaeal lipids record paleosalinity in hypersaline systems. Org. Geochem. 42: 1147-1157 (doi:10.1016/j.orggeochem.2011.06.002) ## salinity salinity,organic compound index,,practical salinity unit,,Paleolimnology; Climate Reconstructions,,mass spectrometry,N,salinity inferred using the linear regression (ACE = 0.35(salinity)-5.4); Turich and Freeman (2011). Archaeal lipids record paleosalinity in hypersaline systems. Org. Geochem. 42: 1147-1157 (doi:10.1016/j.orggeochem.2011.06.002) ## IR6me isomer ratio,,,dimensionless,,Paleolimnology,,mass spectrometry,N,isomer ratio of the sum of 6-methyl isomers to the sum of 5- and 6-methyl isomers: IR6me = (IIa_prime+IIb_prime+IIc_prime+IIIa_prime+IIIb_prime+IIIc_prime)/(IIa+IIb+IIc+IIIa+IIIb+IIIc+IIa_prime+IIb_prime+IIc_prime+IIIa_prime+IIIb'+IIIc_prime); De Jonge et al. (2014) In situ produced branched glycerol dialkyl glycerol tetraethers in suspended particulate matter from the Yenisei River Eastern Siberia. Geochimica Et Cosmochimica Acta 125: 476-491 (doi:10.1016/j.gca.2013.10.031) ## fC fractional cyclization index,,,dimensionless,,Paleolimnology,,mass spectrometry,N,fractional cyclization index of brGDGT: 0.5(((IIIb+IIb+Ib)+2(IIIc+IIc+Ic)+(IIIb_prime+IIb_prime)+2(IIIc_prime+IIc_prime))/(IIIa+IIa+Ia+IIIb+IIb+Ib+IIIc+IIc+Ic+IIIb_prime+IIb_prime+IIIc_prime+IIc_prime)); Martinez-Sosa and Tierney (2019) Lacustrine brGDGT response to microcosm and mesocosm incubations. Organic Geochemistry 127: 12-22 (doi:10.1016/j.orggeochem.2018.10.011) ## archaeol_rerun archaeol,,,dimensionless,,Paleolimnology,,mass spectrometry,N,relative abundance of archaeol; rerun sample (i.e. some fraction of the sample had been previously destructively analyzed this column is the concentration in the rerun for comparison with ex_archaeol to compute R_EA) ## ex_archaeol archaeol,,,dimensionless,,Paleolimnology,,mass spectrometry,N,relative abundance of extended archaeol; rerun sample ## R_EA organic compound index,,,dimensionless,,Paleolimnology,,mass spectrometry,N,index expressing ratio of archaeol to the sum of archaeol and extended archaeol: Vandier et al. (2021) Reappraisal of archaeal C-20-C-25 diether lipid (extended archaeol) origin and use as a biomarker of hypersalinity. Organic Geochemistry 159 (doi:10.1016/j.orggeochem.2021.104276) #------------------------ # Data: # Data lines follow (have no #) # Data line format - tab-delimited text, variable short name as header # Missing_Values: -999999 depth_m age_calyrsBP ID depth_top depth_bottom archaeol isoGDGT-0 isoGDGT-1 isoGDGT-2 isoGDGT-3 isoGDGT-5 isoGDGT-5_prime b-IIIa b-IIIa_prime b-IIIa_3prime b-IIIb b-IIIb_prime b-IIIc bIIIc_prime b-IIa b-IIa_prime b-IIa_3prime b-IIb b-IIb_prime b-IIc b-IIc_prime b-Ia b-Ib b-Ic isoGDGTs_ngg brGDGTs_ngg R_ib IIIa_IIa IIIa_IIa_7Me BIT MBT5Me MAT MAT_2.5th MAT_97.5th MATsoil MATsoil_2.5th MATsoil_97.5th CBTprime pH pHsoil ACE salinity IR6me fC archaeol_rerun ex_archaeol R_EA 0.40 291 GLAD1-GSL00 1B-1H-1 40 41 1244.562 78.924 5.731 11.241 0.335 0.243 0.245 2.522 4.601 21.553 0.324 0.687 0.132 0.122 4.909 14.068 13.985 2.311 2.922 0.106 0.131 16.016 2.107 0.367 96.719 86.862 0.473 0.375 0.870 0.994 0.652 18.975 13.595 24.414 14.719 6.808 22.040 -0.010 8.555 7.211 94.037 284 0.686 0.154 -999999 -999999 -999999 0.70 545 GLAD1-GSL00 1B-1H-1 70 71 1417.554 115.536 6.481 9.551 0.155 1.528 0.111 3.282 6.198 17.285 0.260 0.547 0.048 0.095 7.168 16.453 11.944 3.442 4.211 0.219 0.282 16.718 3.506 0.519 133.363 92.177 0.409 0.401 0.753 0.970 0.595 17.260 11.742 22.582 12.438 4.985 19.981 0.018 8.678 7.267 92.464 280 0.658 0.177 -999999 -999999 -999999 1.00 800 GLAD1-GSL00 1B-1H-1 100 101 1533.197 113.813 4.700 8.024 0.145 2.041 0.117 4.263 8.406 23.772 0.300 0.775 0.059 0.113 9.223 22.157 17.070 4.565 6.028 0.181 0.263 23.907 5.278 0.710 128.840 127.067 0.497 0.404 0.752 0.971 0.621 17.986 12.577 23.583 13.640 6.399 20.555 0.012 8.653 7.256 93.090 281 0.670 0.176 -999999 -999999 -999999 1.30 1046 GLAD1-GSL00 1B-1H-1 130 131 1014.175 145.713 10.020 12.388 0.335 3.024 0.267 4.629 6.829 26.211 0.216 0.531 0.098 0.144 9.182 17.705 20.170 3.559 3.934 0.201 0.291 27.250 3.624 0.631 171.745 125.205 0.422 0.426 0.801 0.956 0.642 18.520 13.103 24.446 14.139 6.985 21.793 -0.135 8.011 6.963 87.437 265 0.622 0.134 -999999 -999999 -999999 1.53 1205 GLAD1-GSL00 1B-1H-2 4 5 1317.927 154.982 9.816 17.575 0.166 1.625 0.096 3.641 6.592 26.885 0.111 0.373 0.072 0.106 7.853 18.289 16.752 3.256 4.380 0.125 0.190 20.607 3.954 0.479 184.259 113.665 0.382 0.391 0.865 0.972 0.627 18.187 12.990 24.549 13.966 6.592 21.069 -0.024 8.498 7.185 89.478 271 0.665 0.155 -999999 -999999 -999999 1.83 1395 GLAD1-GSL00 1B-1H-2 34 35 1485.742 232.471 7.791 10.783 0.227 2.069 0.166 2.706 5.707 27.213 0.113 0.361 0.046 0.158 3.963 12.787 17.120 2.145 3.956 0.066 0.187 19.469 2.165 0.272 253.508 98.432 0.280 0.502 1.052 0.956 0.712 24.207 18.225 29.926 20.129 12.499 27.748 -0.048 8.393 7.137 86.470 262 0.719 0.143 -999999 -999999 -999999 2.24 1653 GLAD1-GSL00 1B-2H-1 16 17 1939.202 139.856 4.637 10.754 0.240 2.822 0.194 3.492 5.689 32.847 0.244 0.463 0.111 0.256 6.124 20.362 27.241 2.581 2.947 0.090 0.179 32.135 2.144 0.309 158.503 137.214 0.464 0.347 0.782 0.960 0.738 21.552 16.074 27.082 17.345 10.023 24.994 -0.141 7.988 6.953 93.273 282 0.703 0.101 0.930 0.070 0.070 2.54 1843 GLAD1-GSL00 1B-2H-1 46 47 1025.766 143.559 6.880 13.038 0.156 1.326 0.115 2.689 5.834 32.444 0.167 0.481 0.072 0.165 5.657 18.637 20.218 2.726 4.158 0.142 0.299 20.752 3.013 0.435 165.073 117.889 0.417 0.351 0.920 0.976 0.683 19.854 14.739 25.908 15.741 8.217 23.110 0.013 8.658 7.258 87.723 266 0.721 0.157 -999999 -999999 -999999 2.82 2020 GLAD1-GSL00 1B-2H-1 74 75 2188.560 161.259 15.223 19.433 0.137 1.694 0.120 3.500 7.766 53.304 0.093 0.505 0.399 0.116 8.464 23.305 30.386 3.563 5.516 0.101 0.129 28.493 4.582 0.486 197.866 170.708 0.463 0.355 1.039 0.977 0.682 19.775 13.981 25.149 15.679 8.265 22.949 -0.029 8.473 7.174 93.137 282 0.698 0.149 -999999 -999999 -999999 3.08 2187 GLAD1-GSL00 1B-2H-1 100 101 927.896 153.443 5.561 14.695 0.198 2.392 0.164 4.166 5.198 61.932 0.019 0.228 0.100 0.241 7.061 22.289 45.158 2.354 2.979 0.029 0.140 48.289 2.203 0.266 176.452 202.652 0.535 0.319 0.957 0.973 0.789 23.273 17.542 28.738 19.209 11.600 26.647 -0.279 7.388 6.680 85.810 261 0.694 0.069 -999999 -999999 -999999 3.38 2387 GLAD1-GSL00 1B-2H-1 130 131 1897.888 351.124 2.364 2.400 0.293 3.097 0.225 5.076 8.220 20.411 0.426 0.753 0.375 0.263 9.239 19.977 14.838 4.558 6.000 0.365 0.501 22.362 4.901 0.861 359.502 119.126 0.249 0.455 0.765 0.954 0.594 17.183 11.870 22.736 12.740 5.017 20.266 -0.001 8.595 7.229 84.388 257 0.641 0.192 -999999 -999999 -999999 3.62 2561 GLAD1-GSL00 1B-2H-2 4 5 739.982 45.026 1.918 4.341 0.099 1.047 0.086 1.750 2.879 13.146 0.100 0.224 0.029 0.117 2.929 6.746 8.657 1.388 1.780 0.081 0.178 9.186 1.437 0.228 52.517 50.854 0.492 0.478 0.970 0.957 0.638 18.751 12.710 23.861 14.265 6.699 21.760 -0.057 8.351 7.118 94.264 285 0.655 0.159 -999999 -999999 -999999 3.91 2779 GLAD1-GSL00 1B-2H-2 33 34 4014.619 145.537 5.848 14.701 0.241 3.435 0.147 2.225 5.426 27.474 0.102 0.330 0.024 0.163 4.883 15.477 16.143 2.413 3.158 0.080 0.402 17.122 2.458 0.464 169.910 98.344 0.367 0.376 0.962 0.929 0.676 19.805 13.887 25.499 15.160 7.477 22.829 0.021 8.691 7.273 96.502 291 0.720 0.159 -999999 -999999 -999999 4.21 3028 GLAD1-GSL00 1B-2H-2 63 64 715.085 107.349 3.709 8.520 0.200 2.392 0.165 3.428 5.851 32.495 0.124 0.392 0.118 0.219 5.864 22.549 23.419 2.589 3.432 0.121 0.414 28.921 2.524 0.420 122.336 132.884 0.521 0.327 0.806 0.965 0.726 21.332 15.880 27.072 17.161 9.514 24.075 -0.060 8.339 7.113 86.947 264 0.729 0.120 -999999 -999999 -999999 4.59 3355 GLAD1-GSL00 1B-2H-2 101 102 1131.403 142.777 9.756 18.752 0.132 1.172 0.084 2.117 4.927 27.745 0.084 0.375 0.121 0.197 5.378 15.773 16.378 2.473 3.409 0.120 0.159 19.264 3.318 0.420 172.673 102.258 0.372 0.333 0.927 0.976 0.695 20.333 14.551 26.249 15.885 8.436 23.442 -0.025 8.491 7.182 88.795 269 0.707 0.156 0.925 0.075 0.075 4.89 3613 GLAD1-GSL00 1B-2H-2 131 132 1235.998 109.119 3.663 7.351 0.165 1.398 0.134 1.717 3.540 18.002 0.121 0.267 0.107 0.151 3.088 9.344 10.836 1.576 2.091 0.083 0.136 11.947 1.518 0.216 121.829 64.739 0.347 0.423 1.000 0.955 0.679 19.939 14.073 25.793 15.325 8.080 22.964 -0.027 8.483 7.178 91.888 278 0.699 0.151 -999999 -999999 -999999 5.12 3797 GLAD1-GSL00 1B-3H-1 13 14 727.276 70.172 2.640 5.531 0.099 0.883 0.073 1.146 2.086 12.699 0.043 0.147 0.031 0.064 2.200 7.350 8.301 1.010 1.292 0.043 0.082 8.381 0.956 0.134 79.399 45.966 0.367 0.338 0.892 0.960 0.683 19.739 14.112 25.552 15.864 8.221 23.621 -0.022 8.506 7.189 91.200 276 0.711 0.134 -999999 -999999 -999999 5.42 4004 GLAD1-GSL00 1B-3H-1 43 44 946.655 174.453 9.908 18.007 0.172 1.459 0.117 2.827 5.450 47.207 0.205 0.545 0.338 0.393 5.667 16.320 28.195 2.494 3.281 0.199 0.310 23.028 2.303 0.373 204.116 139.136 0.405 0.376 1.106 0.973 0.697 20.472 15.062 26.121 15.985 8.731 23.344 -0.073 8.284 7.088 84.439 257 0.692 0.144 -999999 -999999 -999999 5.72 4208 GLAD1-GSL00 1B-3H-1 73 74 1177.023 186.124 4.823 10.609 0.399 3.529 0.366 5.890 7.586 63.013 0.318 0.651 0.360 0.368 9.300 33.859 46.849 3.123 3.668 0.252 0.349 47.856 2.668 0.509 205.849 226.619 0.524 0.312 0.850 0.967 0.733 21.621 15.965 27.596 17.282 9.789 24.498 -0.128 8.045 6.979 86.346 262 0.707 0.094 -999999 -999999 -999999 6.02 4413 GLAD1-GSL00 1B-3H-1 103 104 1137.774 111.304 4.966 8.931 0.262 3.787 0.187 4.091 7.225 33.623 0.251 0.616 0.183 0.258 10.112 28.565 24.742 4.504 4.749 0.260 0.393 32.229 4.460 0.841 129.437 157.102 0.548 0.293 0.709 0.956 0.664 19.463 14.168 24.956 15.088 7.295 23.049 -0.037 8.439 7.158 91.089 276 0.683 0.147 0.973 0.027 0.027 6.32 4622 GLAD1-GSL00 1B-3H-1 133 134 1595.058 192.861 7.551 12.444 0.241 2.197 0.203 4.440 8.425 28.807 0.538 0.942 0.282 0.325 8.640 20.523 18.488 4.326 5.933 0.388 0.564 20.459 4.636 0.817 215.498 128.530 0.374 0.441 0.875 0.966 0.593 17.266 11.532 22.918 12.625 5.198 20.387 0.049 8.813 7.328 89.213 270 0.664 0.202 -999999 -999999 -999999 6.58 4804 GLAD1-GSL00 1B-3H-2 8 9 846.863 195.255 9.048 17.304 0.319 3.190 0.255 4.446 8.285 40.702 0.415 0.763 0.307 0.349 8.999 30.617 29.945 4.287 4.534 0.282 0.418 35.819 4.046 0.933 225.371 175.146 0.437 0.321 0.768 0.965 0.694 20.364 14.601 25.916 15.895 9.008 23.164 -0.031 8.466 7.171 81.264 248 0.706 0.142 -999999 -999999 -999999 6.89 5014 GLAD1-GSL00 1B-3H-2 39 40 1001.537 130.704 8.202 13.080 0.275 3.189 0.238 5.864 8.917 32.946 0.423 0.940 0.121 0.176 10.546 21.878 21.451 4.993 5.943 0.302 0.424 23.713 4.530 0.761 155.688 143.928 0.480 0.456 0.886 0.957 0.572 16.769 11.119 22.100 11.965 4.273 19.637 -0.012 8.548 7.208 88.456 268 0.632 0.174 -999999 -999999 -999999 7.19 5217 GLAD1-GSL00 1B-3H-2 69 70 950.409 111.947 4.247 8.846 0.232 2.260 0.177 2.562 5.021 29.340 0.298 0.495 0.134 0.174 4.806 16.227 19.137 2.476 3.091 0.166 0.238 20.307 2.316 0.490 127.709 107.278 0.457 0.361 0.919 0.956 0.698 20.445 14.710 26.122 16.125 8.495 23.470 -0.032 8.463 7.169 89.462 271 0.707 0.148 -999999 -999999 -999999 7.47 5378 GLAD1-GSL00 1B-3H-2 97 98 2926.722 217.871 14.350 18.479 0.384 5.564 0.316 5.243 9.896 40.276 0.467 0.929 0.219 0.326 9.435 20.075 22.647 4.807 6.256 0.295 0.576 18.451 3.906 0.684 256.963 144.488 0.360 0.513 1.062 0.919 0.538 15.399 9.969 20.889 10.959 3.708 17.962 0.068 8.896 7.366 93.072 281 0.650 0.199 -999999 -999999 -999999 7.77 5537 GLAD1-GSL00 1B-3H-2 127 128 2071.534 195.313 6.151 4.906 0.503 4.905 0.400 7.093 10.784 13.901 0.816 1.315 0.465 0.510 9.048 13.861 14.753 5.642 6.316 0.589 0.789 12.629 4.408 1.034 212.178 103.953 0.329 0.780 0.844 0.916 0.447 16.810 10.848 22.681 12.049 4.783 19.795 0.080 8.949 7.391 91.384 277 0.587 0.249 -999999 -999999 -999999 8.09 5704 GLAD1-GSL00 1B-4H-1 20 21 1132.250 160.413 4.907 5.027 0.389 4.011 0.322 5.480 8.226 12.156 0.468 0.821 0.115 0.218 7.785 12.469 13.331 4.514 5.481 0.358 0.594 10.974 3.855 0.748 175.070 87.593 0.333 0.677 0.770 0.918 0.462 13.281 7.715 19.373 8.529 1.599 16.271 0.071 8.910 7.373 87.590 266 0.598 0.232 0.757 0.243 0.243 8.41 5872 GLAD1-GSL00 1B-4H-1 52 53 1699.373 148.758 2.702 2.263 0.353 4.049 0.289 5.870 9.179 12.376 0.502 0.826 0.081 0.199 9.033 14.539 15.314 5.309 5.227 0.352 0.690 12.377 4.634 0.839 158.414 97.346 0.381 0.638 0.705 0.926 0.465 15.518 10.256 20.940 10.799 3.695 18.450 0.062 8.872 7.355 91.951 278 0.592 0.227 -999999 -999999 -999999 8.71 6035 GLAD1-GSL00 1B-4H-1 82 83 1257.165 131.153 5.003 4.099 0.330 3.283 0.299 5.018 8.169 16.651 0.427 0.727 0.094 0.201 7.341 13.442 17.981 4.318 5.398 0.329 0.631 13.922 3.873 0.764 144.166 99.285 0.408 0.635 0.770 0.936 0.522 15.135 9.434 20.542 10.182 2.923 17.787 0.048 8.808 7.326 90.553 274 0.620 0.218 -999999 -999999 -999999 9.06 6240 GLAD1-GSL00 1B-4H-1 117 118 2318.452 149.956 1.962 1.119 0.285 2.611 0.207 4.646 7.577 4.888 0.406 0.670 0.035 0.156 7.189 10.411 8.091 4.006 5.196 0.304 0.749 8.446 3.685 0.704 156.140 67.159 0.301 0.694 0.666 0.936 0.443 12.434 6.667 17.529 7.944 0.734 15.935 0.099 9.030 7.427 93.925 284 0.599 0.247 0.699 0.301 0.301 9.33 6396 GLAD1-GSL00 1B-4H-1 144 145 2228.456 130.958 2.523 1.957 0.404 4.160 0.331 4.745 8.235 11.882 0.520 0.740 0.128 0.305 7.143 13.183 15.224 4.218 5.116 0.418 0.798 13.095 3.791 0.875 140.332 90.416 0.392 0.639 0.699 0.918 0.518 15.132 9.640 20.535 10.070 3.036 17.706 0.069 8.898 7.367 94.450 285 0.623 0.232 -999999 -999999 -999999 9.66 6592 GLAD1-GSL00 1B-4H-2 27 28 2548.995 163.427 2.480 1.303 0.370 4.148 0.293 6.556 10.791 4.901 0.634 1.113 0.106 0.322 10.378 15.368 11.601 5.700 7.842 0.460 1.125 11.965 5.429 1.182 172.021 95.472 0.357 0.674 0.596 0.930 0.446 12.791 6.848 18.632 8.013 1.084 15.174 0.116 9.105 7.461 93.975 284 0.605 0.257 -999999 -999999 -999999 9.96 6776 GLAD1-GSL00 1B-4H-2 57 58 2506.559 144.798 1.928 1.537 0.252 3.556 0.221 5.730 9.432 11.494 0.513 0.829 0.057 0.189 10.986 18.010 16.304 5.899 7.753 0.387 0.886 17.525 6.802 1.216 152.291 114.011 0.428 0.523 0.588 0.945 0.526 15.138 9.399 20.815 10.413 3.263 18.416 0.049 8.813 7.328 94.539 286 0.611 0.232 0.933 0.067 0.067 10.26 6961 GLAD1-GSL00 1B-4H-2 87 88 4838.982 101.701 3.096 3.779 0.449 6.118 0.380 5.094 9.286 25.179 0.381 0.791 0.081 0.233 8.676 18.907 16.431 4.758 6.384 0.318 0.853 20.974 4.523 0.906 115.523 123.776 0.517 0.521 0.899 0.911 0.584 17.125 11.484 22.830 12.239 5.138 20.374 0.032 8.737 7.294 97.942 295 0.654 0.200 -999999 -999999 -999999 10.66 7206 GLAD1-GSL00 1B-4H-2 127 128 1620.046 188.844 2.540 1.197 0.290 3.301 0.229 6.252 10.205 7.604 0.620 0.940 0.080 0.276 10.235 16.126 11.883 5.603 7.844 0.520 1.103 14.433 6.026 1.203 196.400 100.954 0.340 0.624 0.629 0.945 0.489 14.073 8.276 19.641 9.375 1.709 16.530 0.086 8.975 7.402 89.560 271 0.610 0.248 -999999 -999999 -999999